von Konrat, M.; Braggins, J.; Asakawa, U.; Toyota, M. 2006: Recognition of Frullania congesta: a case study to present a species concept and a synthesis of significant taxonomic characters for the large liverwort genus Frullania (Frullaniaceae). Journal of the Hattori Botanical Laboratory 100: 553–576.
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von Konrat, M.; Braggins, J.; Asakawa, U.; Toyota, M. 2006: Recognition of Frullania congesta: a case study to present a species concept and a synthesis of significant taxonomic characters for the large liverwort genus Frullania (Frullaniaceae). Journal of the Hattori Botanical Laboratory 100: 553–576.