Engel, J.J.; Schuster, R. M. 1985 ("1984"): An overview and evaluation of the genera of Geocalycaceae subfamily Lophocoleoideae (Hepaticae). Nova Hedwigia 39: 385–445.

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Engel, J.J.; Schuster, R. M. 1985 ("1984"): An overview and evaluation of the genera of Geocalycaceae subfamily Lophocoleoideae (Hepaticae). Nova Hedwigia 39: 385–445.