Murakami, N.; Nogami, S.; Watanabe, M.; Iwatsuki, K. 1999: Phylogeny of Aspleniaceae inferred from rbcL nucleotide sequences. American Fern Journal 89: 232–243.

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Murakami, N.; Nogami, S.; Watanabe, M.; Iwatsuki, K. 1999: Phylogeny of Aspleniaceae inferred from rbcL nucleotide sequences. American Fern Journal 89: 232–243.